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The persistent inter‐model spread in the response of global‐mean surface temperature to increased CO₂(known as the “Equilibrium Climate Sensitivity,” or “ECS”) is a crucial problem across model generations. This work examines the influence of the models' present‐day atmospheric circulation climatologies, and the accompanying climatological cloud radiative effects, in explaining that spread. We analyze the Coupled Model Intercomparison Project Phase 6 (CMIP6) models and find that they simulate a more poleward, and thus more realistic, edge of the Hadley cell (HC) in the Southern Hemisphere than the CMIP5 models, although the climatological shortwave cloud radiative effects are similar in the two generations of models. A few CMIP5 models with extreme equatorward biases in the HC edge exhibited high ECS due to strong Southern midlatitude shortwave cloud radiative warming in response to climate change, suggesting an ECS dependence on HC position. We find that such constraint no longer holds for the CMIP6 models, due to the absence of models with extreme HC climatologies. In spite of this, however, the CMIP6 models show an increased spread in ECS, with more models in the high ECS range. In addition, an improved representation of the climatological jet dynamics does not lead to a new emergent constraint in the CMIP6 models either.

Recent analyses of global climate models suggest that uncertainty in the coupling between midlatitude clouds and the atmospheric circulation contributes to uncertainty in climate sensitivity. However, the reasons behind model differences in the cloud–circulation coupling have remained unclear. Here, we use a global climate model in an idealized aquaplanet setup to show that the Southern Hemisphere climatological circulation, which in many models is biased equatorward, contributes to the model differences in the cloud–circulation coupling. For the same poleward shift of the Hadley cell (HC) edge, models with narrower climatological HCs exhibit stronger midlatitude cloud-induced shortwave warming than models with wider climatological HCs. This cloud-induced radiative warming results predominantly from a subsidence warming that decreases cloud fraction and is stronger for narrower HCs because of a larger meridional gradient in the vertical velocity. A comparison of our aquaplanet results with comprehensive climate models suggests that about half of the model uncertainty in the midlatitude cloud–circulation coupling stems from this impact of the circulation on the large-scale temperature structure of the atmosphere, and thus could be removed by improving the climatological circulation in models. This illustrates how understanding of large-scale dynamics can help reduce uncertainty in clouds and their response to climate change.